Publications

Anderson, C. J., Weand, M. P., Alexander, H. D., Bretfeld, M., Green, N. 2025. Variations in stand structure, composition, and fuelbeds drive prescribed fire behavior during mountain longleaf pine restoration. Forest Ecology and Management 575: 122372. Download.

Webb, E. E., Alexander, H. D., Loranty, M. M., Talucci, A. C., Lichstein, J. W. 2024. Controls over fire characteristics in Siberian larch forests. Ecosystems. https://doi.org/10.1007/s10021-024-00927-8 Download.

Lazzaro, L. G., Alexander, H. D., Cannon, J. B., Aspinwall, M. J. 2024. Overstory and fuel traits drive moisture dynamics of mesophytic and pyrophytic leaf litter and 10-hour woody debris fuels in a mixed longleaf pine-hardwood woodland. Fire Ecology 20 (63). Download.

Willis, J., Alexander, H. D., Milton, T. 2024. Cone and fruit impacts on understory flammability depend on traits and forest floor coverage. Fire Ecology 20 (52). Download.

Alexander, H. D., Paulson, A. K., Loranty, M., Natali, S., Mack, M. C., Peña III, H., Davydov, S., Zimov, N. 2024. Linking post-fire tree density to carbon storage in high-latitude cajander larch (Larix cajanderi) forests of Far Northeastern Siberia. Ecosystems. Download.

Berner, L.T., Orndahl, K. M., Rose, M., Tamstorf, M., Arndal, M. F., Yan, D., Alexander, H. D., +46 authors. 2024. The Arctic Plant Aboveground Biomass Synthesis Dataset. Scientific Data. 11: 305. Download.

Loranty, M. M., Alexander, H. D., Davydov, Kholodov, A. L., S. P., Kropp, H., M., Mack, Natali, S. M. C., Zimov, N. S. 2024. Winter soil temperature varies with canopy cover in Siberian larch forests. Environmental Research Letters 19(5): 054013. Download.

Webb, E. E., Alexander, H. D., Paulson, A. K., Loranty, M. M., DeMarco, J., Talucci, A. C., Spektor, V., Zimov, N. Lichstein, J. W. 2024. Carbon loss and tree mortality following fire in Siberian larch forests. Geophysical Research Letters 51(1): e2023GL105216. Download.

Scavotto, N. A., Siegert, C. M., Alexander, H. D., Varner, J. M. 2024. Bark and crown morphology drive differences in rainwater distribution in an upland oak forest. Forest Ecology and Management 553: 121642. Download.

Borth, E., Alexander, H. D., Zimov, N., McEwan, R. 2023. Seed sources and safe sites as drivers of Larix cajanderi regeneration following fire in the Siberian Arctic. Ecosphere. 14:e4617. Download.

Cabrera, S. M., Alexander, H. D., Willis, J. L., Anderson, C. J. 2023. Midstory removal of encroaching species has minimal impacts on fuels and fire behavior regardless of burn season in a degraded pine-oak mixture. Forest Ecology and Management 544: 121157. Download.

Goldsmith, C., Alexander, H. D., Granger, J., Siegert, C. 2023. Impacts of Microstegium vimineum on performance of three hardwood species under different growth environments. Forest Ecology and Management 539: 120984. Download.

Bendavid, N. S., Alexander, H. D., Davydov, S. P., Kropp, H., Mack, M. C., Natali, S. M., Spawn-Lee, S. A., Zimov, N., Loranty, M. M. 2023. Shrubs compensate for tree leaf area variation and influence vegetation indices in post-fire Siberian larch forests. JGR-Biogeosciences. 128(3): e2022JG007107. Download

Willis, J. L., Renninger, H. J., Schnake, D. K, Alexander, H. D. 2023. Xerophytic hardwood retention promotes competition over facilitation in longleaf pine woodlands in the absence of fire. Forest Ecology and Management 531: 120732. Download

Hewitt, R., Alexander, H. D., *Izbicki, B., Loranty, M. A., Natali, S. M., Walker, X. J., Mack, M. C. 2022. Increasing tree density accelerates stand-level nitrogen cycling at the taiga-tundra ecotone in northeastern Siberia. Ecosphere 13(7): e4175. Download

Hewitt, R., Alexander, H. D., Miller, S. N., Mack, M. C. 2022. Root-associated fungi not tree density drives nitrogen cycling at the tundra-taiga ecotone. Journal of Ecology 00: 1-13. Download

Babl-Plauche, E., Alexander, H. D., Willis, J., Berry, A. 2022. Mesophication of upland oak forests: implications of species-specific differences in leaf litter decomposition rates and fuelbed composition. Forest Ecology and Management 512: 120141. Download

Boggess, C. M., Baruzzi, C., Alexander, H. D., Strickland, B., Lashley, M. 2022. Exposure to fire affects acorn removal by altering consumer preference. Forest Ecology and Management 508: 120044 Download

Talucci, A. C., Loranty, M., Alexander, H. D. 2022. Spatial patterns of unburned refugia in Siberian larch forests during the exceptional 2020 fire season. Global Ecology and Biogeography 00: 1-15. Download

Talucci, A. C., Loranty, M., Alexander, H. D. 2022. Siberian taiga and tundra fire regimes from 2001-2020. Environmental Research Letters 17:025001. Download

Loranty, M., Alexander, H. D., Kropp, H., Talucci, A. C., Webb, E. E. 2021. Siberian ecosystems as drivers of cryospheric climate feedbacks in the terrestrial Arctic. Frontiers in Climate 3: 730943. Download

Boggess, C., Mason, D., Alexander, H. D., Strickland, B., Lashley, M. 2021. Facultative seed predators drive community-level indirect effects of mast seeding. Forest Ecology and Management 502: 119713. Download

Baltzer, J. L., Day, N. J., Walker, X. J., Mack, M. C., Alexander, H. D., Arsenault, D., Barnes, J., Boucher, Y., Bourgeau-Chavez, L., Brown, C. D., Carrière, S., Howard, B., Greene, D., Gauthier, S., Parisien, M. A., Reid, K. A., Rogers, B. M., Roland, C., Sirois, L., Stehn, S., Thompson, D., Turetsky, M. R., Veraverbeke, S., Whitman, E. J., Yang, J., Johnstone, J. F. 2021. Increasing fire and the decline of fire adapted black spruce in the boreal forest. Proceedings of the National Academy of Sciences 118(45): e2024872118. Download

Fan, J., Song, A., Dong, L., Alexander, H. D., Yang, S., Cheng, N., Pitchford, J. L. 2021. Fire effects on post-invasion spread of Chinese tallow (Triadica sebifera) in wet pine flatwood ecosystems in the southeastern United States. Forest Ecology and Management 500: 119658. Download

Walker, X., Alexander, H. D., Berner, L. T., Boyd, M., Loranty, M., Mack, M. C. 2021. Positive response of tree productivity to warming is reversed by increased tree density at Arctic treeline. Canadian Journal of Forest Research. 51(9): 1323 – 1338. Download

Paulson, A., *Peña III, H., Alexander, H. D., Davydov, S., Loranty, M. M., Mack, M. C., Natali, S. M. 2021. Understory plant diversity and composition across a post-fire tree density gradient in a Siberian Arctic boreal forest. Canadian Journal of Forest Research 51(5): 720-731. Download

Mack, M. C., Walker, X. J., Johnstone, J. F., Alexander, H. D., Melvin, A. M., Jean, M., Miller, S. N. 2021. Carbon loss from boreal forest wildfires offset by increased dominance of deciduous trees. Science 372 (6539): 280-283. Download

Alexander, H. D., Siegert, C., Brewer, S., Kreye, J., Lashley, M., *McDaniel, J., #Paulson, A., Renninger, H., Varner, M. 2021. Mesophication of oak landscapes: Evidence, knowledge gaps, and future research. BioScience 1-12. Download

McDaniel, J. K., Alexander, H. D., Siegert, C. M., Lashley, M. A. 2021. Shifting tree species composition of upland oak forests alters leaf litter structure, moisture, and flammability. Forest Ecology and Management 482: 118860. Download

Nation, R., Alexander, H. D., *McDaniel, J. K., Denny, G., #Paulson, A. 2021. Impacts of increasing fine fuel loads on acorn germination and early growth of oak seedlings. Fire Ecology 17(1): 1 – 13. Download

Talucci, A. C., Forbath, E., Kropp, H., Alexander, H. D., #Paulson, A. K., Zimov, N. S., Loranty, M. M. 2020. Evaluating post-fire vegetation recovery in Cajander larch forests in Northeastern Siberia using UAV derived vegetation indices. Remote Sensing 12(18): 2970. Download

Hanberry, B., Alexander, H. D., Bragg, D. C. 2020. Open forest ecosystems: an excluded state with conservation costs. Forest Ecology and Management 472: 118256. Download

Arney, R., *Shepherd, A., Alexander, H. D., Rahman, A. 2020. Soil carbon and nitrogen storage in natural and prop-scarred Thalassia testudinum seagrass meadows. Estuarine, Coastal, and Shelf Science. Download

Izbicki, B. J., Alexander, H. D., Frey, B. R., McEwan, R. W., Berry, A. I. 2020. Prescribed fire and natural canopy gap disturbances: impacts on upland oak regeneration. Forest Ecology and Management 465: 118107. Download

Thomas, H.J.D., Bjorkman, A.D., Myers-Smith, I.H., Elmendorf, S.C., Kattge, J., Diaz, S., Vellend, M., Blok, D., Cornelissen, J.H.C., Forbes, B.C., Henry, G.H.R., Hollister, R.D., Normand, S., Prevéy, J.S., Rixen, C., Schaepman-Strub, G., Wilmking, M., Wipf, S., Cornwell, W.K., Beck, P.S.A., Georges, D., Goetz, S.J., Guay, K.C., Rüger, N., Soudzilovskaia, N.A., Spasojevic, M.J., Alatalo, J.M. (many authors, including Alexander, H. D.). 2020. Global plant trait relationships extend to the climatic extremes of the tundra biome. Nature Communications 11, 1351. Download

Babl, E. K., Alexander, H. D., Siegert, C. M., Willis, J. W. 2020. Could canopy, bark, and leaf litter traits of encroaching non-oak species influence future flammability of upland oak forests? Forest Ecology and Management. Download

Siegert, C. M., Drotar, N., Alexander, H. D. 2019. Spatial and temporal variability of throughfall in an upland oak-hickory forest. Geosciences 9 (405):1–13. Download

Kropp, H., Loranty, M., Natali, S., Kholodov, A., Alexander, H. D., Zimov, N., Mack, M., Spawn, S. 2019. Tree density influences ecohydrological drivers of plant–water relations in a larch boreal forest in Siberia. Ecohydrology 12(7):e2132. Download

Watson, P., Alexander, H. D. 2019. Effects of shrub encroachment and removal on grassland recovery in South Texas Coastal Prairies. Rangeland Ecology and Management 72:275–282. Download

Loranty, M. M., Kropp, H., Natali, S. M., Alexander, H. D., Mack, M. C., Zimov, N., Davydov, S. P. 2018. Vegetation indices do not capture forest cover variation in upland Siberian larch forests. Remote Sensing 10:1686. Download

Bjorkman, A. D., Myers-Smith, I. H., Elmendorf, S. C., Normand, S., Thomas, H., et al. (many authors, including Alexander, H. D.). 2018. Tundra Trait Team: A database of plant traits spanning the tundra biome. Global Ecology and Biogeography 2018:1–10. Download

Bjorkman, A. D., Myers-Smith, I. H., Elmendorf, S. C., Rüger, N., Normand, S., et al. (many authors, including Alexander, H. D.). 2018. Plant functional trait change across the tundra biome and over three decades of warming. Nature 562:57–62. Download

Ludwig, S., Alexander, H. D., Kielland, K., Mann, P., Natali, S. M., Ruess, R. 2018. Fire severity effects on nutrient dynamics and microbial activities in a Siberian larch forest. Global Change Biology 2018:1–12. Download

Alexander, H. D., Natali, S., Loranty, M., Mack, M. C., Ludwig, S., Davydov, S., Zimov, N. 2018. Soil burn severity impacts on larch recruitment on permafrost soils of far northeastern Siberia. Forest Ecology and Management 417: 144-153. Download

Alexander, H. D., M. C. Mack. 2017. Gap regeneration within mature deciduous forests of Interior Alaska: implications for future forest successional dynamics. Forest Ecology and Management 396(2017): 35-43. Download

Arthur, M. A., B. A. Blankenship, A. Schörgendorfer, H. D. Alexander. 2017. Alterations to the fuel bed after single and repeated prescribed fires in an Appalachian hardwood forest. Forest Ecology and Management 403:126-136. Download

Jean, M., J. Johnstone, M. Mack, H.D. Alexander. 2017. Patterns of bryophyte succession in a 160 year chronosequence in deciduous and coniferous forests of boreal Alaska. Canadian Journal of Forest Research 47(8):1021-1032. Download

Kropp, H., M. Loranty, H. D. Alexander, L. Berner, S. Natali, S. Spawn. 2017. Hydrologic suppression of transpiration and stomatal conductance in a Siberian Arctic boreal forest. Journal of Geophysical Research: Biogeosciences 122: 487-497. Download

Webb, E. E., K. Heard, S. Natali, A. Bunn, H. D. Alexander, L. Berner, A. Kholodov, M. Loranty, J. Schade, V. Spektor, N. Zimov. 2017. Variability in above and belowground carbon stocks in a Siberian larch watershed. Biogeosciences 14:4270-4294. Download

Abbott, B.W., J. B. Jones, E. A. G. Schuur, F. S. Chapin III, W. B. Bowden, M. S. Bret-Harte, H. E. Epstein, M. D. Flannigan, T. K. Harms, T. N. Hollingsworth, M. C. Mack, A. D. McGuire, S. M. Natali, A. V. Rocha, S. E. Tank, M. R. Turetsky, J. E. Vonk, K. P. Wickland, G. R. Aiken, H. D. Alexander, etal. 2016. Biomass offsets little or none of permafrost carbon release from soils, streams, and wildfire: an expert assessment. Environmental Research Letters 11(3): 1-13. Download

Alexander, H. D. and Mack, M. C. 2016. A canopy shift in interior Alaskan boreal forests: consequences for above and belowground carbon and nitrogen pools during post fire succession. Ecosystems 19(1):98-114. Download

Alexander, H. D., Moczygemba, J., and Dick, K. 2016. Growth and survival of thornscrub forest seedlings in response to restoration strategies aimed at alleviating abiotic and biotic stressors. Journal of Arid Environments 124: 180-188. Download

Dick, K., H. D. Alexander, J. D. Moczygemba. 2016. Use of shelter tubes, grass-specific herbicide, and herbivore exclosures to reduce stressors and improve restoration of semi-arid thornscrub forests. Restoration Ecology 24(6):785-793. Download

Loranty, M., W. Liberman-Cribbin, L. T. Berner, H. D. Alexander, S. J. Goetz, A. L. Kholodov, S. N. Natali. 2016. Spatial variation in vegetation productivity trends, fire distrubance, and soil carbon acros arctic-boreal permafrost ecosystems. Environmental Research Letters 11(9):1-14. Download

Arthur, M. A., Blankenship, B.A., Schorgendorfer, A., Carmean, G., Loftis, D. L., and Alexander, H. D., 2015. Stand structure and tree health following repeated prescribed fire in an Appalachian hardwood forest. Forest Ecology and Management 340: 46-61. Download

Berner, L. T., H.D. Alexander, M.M. Loranty, P. Ganzlin, M.C. Mack, S.J. Goetz. 2015. Biomass allometry for alder, dwarf birch, and willow in boreal forest and tundra ecosystems of northeastern Siberia and northern Alaska. Forest Ecology and Management 337:110-118. Download

Broderick, D.E., K.E. Frey, J. Rogan, H.D. Alexander. 2015. Estimating belowground carbon stocks in a permafrost watershed in Northeast Siberia by linking field measurements to Landsat – 5 TM and Worldview – 2 data. GIS and Remote Sensing 52(2):1-25. Download

Lei, J., B.K. Wylie, D.R. Nossov, B. Peterson, H.D. Alexander, M.C. Mack, J. Rover, M.P. Waldrop, J.W. McFarland, X. Chen, N.J. Pastick. 2015. Spatially explicit estimation of aboveground boreal forest biomass in the Yukon River Basin, Alaska. International Journal of Remote Sensing 36(4):939-953. Download

Alexander, H. D., M. A. Arthur. 2014. Increasing red maple leaf litter alters decomposition rates and nitrogen cycling in historically oak – dominated forests of the eastern U.S. Ecosystems 17:1371-1383. Download

McLauchlan, K.K., P.E. Higuera, D.G. Gavin, S.P. Perakis, M.M. Mack, H.D. Alexander, J. Battles, F. Biondi, B. Buma, D. Colombaroli, S. Enders, D.R. Engstrom, F.S. Hu, J. Marlon, J. Marshall, M. McGlone, J. Morris, L. Nave, B. Shuman, E. Smithwick, D. Urrego, D. Wardle, C. Williams, J.J. Williams. 2014. Reconstructing disturbances and their biogeochemical consequences over multiple timescales. Bioscience 64(2):105-116. Download

Alexander, H. D., M.C. Mack, S. Goetz, P.S.A. Beck, F. Belshe. 2012. Implications of increased deciduous cover on stand structure and aboveground carbon pools of Alaskan boreal forests. Ecosphere 3(5):45. Downlaod

Alexander, H. D., M.C. Mack, S. Goetz, P.S.A. Beck, M. Loranty, K. Earl, S. Zimov, S. Davydov, C.C. Thompson. 2012. Carbon accumulation patterns during post– fire succession in Cajander larch (Larix cajanderi) forests of Siberia. Ecosystems 15(7): 1065-1082. Download

Arthur, M. A., H.D. Alexander, D. Dey, C.J. Schweitzer, D.L. Loftis. 2012. Refining the oak – fire hypothesis for management of oak – dominated forests of the eastern United States. Journal of Forestry 110(5): 257-266. Download

Berner, L. T., P.S.A. Beck, M.M. Loranty, H.D. Alexander, M.C. Mack, S.J. Goetz. 2012. Cajander larch (Larix cajanderi) biomass distribution, fire regime and post – fire recovery in northeastern Siberia. Biogeosciences 9: 3943-3959. Download

Beck, P. S. A.,S. Goetz, M.C. Mack, H.D. Alexander, J. Yufang, J.T. Randerson, M. Loranty. 2011. The impacts and implications of an intensifying fire regime on boreal forest composition and albedo. Global Change Biology 17: 285-2866. Download

Alexander, H. D., M. A. Arthur. 2010. Implications of a predicted shift from upland oaks to red maple on forest hydrology and nutrient availability. Canadian Journal of Forest Research 40(4): 716-726. Download

Alexander, H. D., M. A. Arthur. 2009. Foliar morphology and chemistry of upland oaks, red maple, and sassafras seedlings in response to single and repeated prescribed fires. Canadian Journal of Forest Research 39: 740-754. Download

Alexander, H. D., M. A. Arthur, D. L. Loftis, S. R. Green. 2008. Survival and growth of upland oak and co – occurring competitor seedlings following single and repeated prescribed fires. Forest Ecology and Management 256: 1021-1030. Download

Forbes, M., H. D. Alexander, K. H. Dunton. 2008. The effects of pulsed riverine versus non – pulsed wastewater inputs of freshwater on plant community structure in a semi – arid salt marsh. Wetlands 28(4): 984-994. Download

Alexander, H. D., K. H. Dunton. 2006. Wastewater effluent as an alternative freshwater source in a hypersaline salt marsh: Impacts on salinity, inorganic nitrogen, and emergent vegetation. Journal of Coastal Research 22(2): 377-392. Download

Alexander, H. D., K. H. Dunton. 2002. Freshwater inundation effects on emergent vegetation of a hypersaline salt marsh. Estuaries 25(6B): 1426-1435. Download